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The territories of mature females were always enclosed within the territory of a single, mature male. Social and mating systems of this species are based upon female territoriality where polygyny results from males defending females occupying a certain territory (female-defense polygyny) (Gladstone 1987).Neither parent guards the eggs which are laid in a nest located in the female's territory (Gladstone 1987).The eggs and larvae of this species are unpalatable to other species (Gladstone 1987).Larvae may have high survivorship as a result (Gladstone and Westoby 1988).

are known for their general lack of characters, and are generally differentiated on the basis of colour differences.

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